The tracks left by organisms are among the most difficult of fossils to interpret. But just such evidence puts debate about the origins of four-limbed vertebrates (which include ourselves) on a changed footing.
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Janvier, P., Clément, G. Muddy tetrapod origins. Nature 463, 40–41 (2010). https://doi.org/10.1038/463040a
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DOI: https://doi.org/10.1038/463040a
Erik Schlenker
What geological factors could lead to this particular ghost range? In general, I assume a climate change could lead to a ghost range if fossilization happens less readily in one climate over another, but I am not sure if this would apply to this time span